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HOWLish : a CNN for automated wolf howl detection
ID Campos, Rafael (Author), ID Krofel, Miha (Author), ID Rio-Maior, Helena (Author), ID Renna, Francesco (Author)

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Abstract
Automated sound-event detection is crucial for large-scale passive acoustic monitoring of wildlife, but the availability of ready-to-use tools is narrow across taxa. Machine learning is currently the state-of-the-art framework for developing sound-event detection tools tailored to specific wildlife calls. Gray wolves (Canis lupus), a species with intricate management necessities, howl spontaneously for long-distance intra- and inter-pack communication, which makes them a prime target for passive acoustic monitoring. Yet, there is currently no pre-trained, open-access tool that allows reliable automated detection of wolf howls in recorded soundscapes. We collected 50 137 h of soundscape data, where we manually labeled 841 unique howling events. We used this dataset to fine-tune VGGish—a convolutional neural network trained for audio classification—effectively retraining it for wolf howl detection. HOWLish correctly classified 77% of the wolf howling examples present on our test set, with a false positive rate of 1.74%; still, precision was low (0.006) granted extreme class imbalance (7124:1). During field tests, HOWLish retrieved 81.3% of the observed howling events while offering a 15-fold reduction in operator time when compared to fully manual detection. This work establishes the baseline for open-access automated wolf howl detection. HOWLish facilitates remote sensing of wild wolf populations, offering new opportunities in non-invasive large-scale monitoring and communication research of wolves. The knowledge gap we addressed here spans across many soniferous taxa, to which our approach also tallies.

Language:English
Keywords:bioacoustics, deep learning, howling, monitoring, wolf
Work type:Article
Typology:1.01 - Original Scientific Article
Organization:BF - Biotechnical Faculty
Publication status:Published
Publication version:Version of Record
Year:2026
Number of pages:Str. 58-70
Numbering:Vol. 12, iss. 1
PID:20.500.12556/RUL-180425 This link opens in a new window
UDC:[599.744.111.1:591.582]:004.85
ISSN on article:2056-3485
DOI:10.1002/rse2.70024 This link opens in a new window
COBISS.SI-ID:250212611 This link opens in a new window
Publication date in RUL:09.03.2026
Views:324
Downloads:169
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Record is a part of a journal

Title:Remote sensing in ecology and conservation
Shortened title:Remote sens. ecol. conserv.
Publisher:Zoological Society of London
ISSN:2056-3485
COBISS.SI-ID:525560345 This link opens in a new window

Licences

License:CC BY-NC 4.0, Creative Commons Attribution-NonCommercial 4.0 International
Link:http://creativecommons.org/licenses/by-nc/4.0/
Description:A creative commons license that bans commercial use, but the users don’t have to license their derivative works on the same terms.

Secondary language

Language:Slovenian
Keywords:bioakustika, strojno učenje, oglašanje, monitoring, volk, Canis lupus

Projects

Funder:FCT - Fundação para a Ciência e a Tecnologia, I.P.
Project number:2021.08079.BD
Name:Listening for Wolf Conservation: Deep Learning for Automated Howl Recognition and Classification.

Funder:ARIS - Slovenian Research and Innovation Agency
Project number:J1-50013
Name:ExtremePredator: Odkrivanje ekološke vloge vrhovnih plenilcev v ekstremnih okoljih

Funder:ARIS - Slovenian Research and Innovation Agency
Project number:P4-0059
Name:Gozd, gozdarstvo in obnovljivi gozdni viri

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